Abstract: An in-line roller skate and shoe combination includes a shoe having a sole with at least one recess therein opening downward to the bottom face. The bottom face of the sole is generally flat and free of protrusions to permit walking when the wheel support frame is not engaged. A wheel support frame has at least two in-line wheels rotatably secured to a base of the frame. The recess forms a longitudinal track running from a region of the sole adjacent a toe end of the sole towards and opening out of a heel end of the sole and continuing, curving upwards along the back of the shoe. A wheel support frame assembly has wheels residing rotatably, each within a separate wheel housing, with each wheel housing being linked, preferably by a flexible wire. An upper portion of each wheel housing is shaped to conform to the track so as to be able to ride along said track while being supportingly retained therein.