Abstract: Apparatus is provided for optically measuring scalar irradiance or incident flux of radiant energy and for optically measuring naturally occurring chlorophyll fluorescence or upwelling radiance in a parcel of water in a natural setting. From a comparison of the two measurements, rate of primary photosynthetic production is calculated by appropriately programmed computer means. Readout means are provided to indicate the rate of primary production. Further, concentrations of chlorophyll that generate the primary production are also determined by means of the apparatus of this invention. The method of this invention, employing measurement instruments of the type disclosed, enables the determination of concentrations of chlorophyll and primary production in parcels of water.