Abstract: Signal charges are read out from light-receiving portions during a vertical blanking period by applying a read-out voltage pulse to read-out electrodes that are provided separately from vertical transfer electrodes, while signal charges are prevented from leaking from the light-receiving portions during a vertical scanning period by applying a negative voltage to the read-out electrodes. The read-out voltage pulse also is applied to vertical transfer electrodes, whereby signal charges are read out from the light-receiving portions more efficiently. Signal charges are read out from light-receiving portions in a predetermined region by applying a read-out voltage pulse to read-out electrodes while applying a negative voltage to a part of the vertical transfer electrodes.