Abstract: A Cucumis sativus var. sativus plant tolerant to Tomato Leaf Curl New Delhi Virus (ToLCNDV) includes in its genome the combination of a first quantitative trait locus (QTL) QTL1 on chromosome 1 and a second QTL, QTL2 on chromosome 2, at least one of QTL1 and QTL2 being homozygous, wherein said combination confers to the plant tolerance to ToLCNDV and said QTLs on chromosomes 1 and 2 are present in the genome of the seeds of plant TOCUR6080, NCIMB accession number 43427. The QTL are preferably characterized by defined alleles of different SNPs on chromosomes 1 and 2. Parts of these plants have ToLCNDV tolerance phenotype, as well as progeny, and can be used for introgressing the tolerance in another genetic background, with different methods for obtaining cucumber plants or seeds with increased tolerance to ToLCNDV and different markers linked to the QTLs used to confer tolerance phenotype.