Abstract: An aerosol-generating article is provided, including an aerosol-generating substrate, the aerosol-generating substrate formed of a homogenised plant material, including: between 1 percent by weight and 65 percent by weight of non-tobacco plant particles, on a dry weight basis; between 15 percent by weight and 55 percent by weight of aerosol former, on a dry weight basis; between 5 percent by weight and 10 percent by weight of cellulose ether, on a dry weight basis; and between 5 percent by weight and 50 percent by weight of additional cellulose, on a dry weight basis, in which the additional cellulose is in a form of isolated cellulose and is not derived from the non-tobacco plant particles, and in which a ratio of additional cellulose to cellulose ether in the homogenised plant material is at least 2.

Abstract: An aerosol-forming substrate is provided, including: a) one or both of cellulose and cellulose derivatives; b) an aerosol-former present in an amount of 20 weight percent to 58 percent on a dry weight basis based on a total amount of the aerosol-forming substrate; c) from 0 weight percent to 5 weight percent of tobacco on a dry weight basis based on the total amount of the aerosol-forming substrate; d) a nitrogen-containing nucleophilic compound; and e) a disaccharide. An aerosol-generating article including a substrate portion containing the aerosol-forming substrate is also provided. An aerosol-generating system is also provided.