Zoysiagrass plant named `Victoria`
A new and distinct variety of Zoysiagrass which has a longer growing season and green color retention in winter and less anthocyanin than the variety `El Toro`.
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The present invention relates to a new and distinct variety of Zoysia sp., which is a member of the Gramineae family. The parents of `Victoria`, also referred to herein as `Z88-14` are native to Asia.
The new variety is the result of crossing the Zoysiagrass variety `El Toro` (U.S. Plant Pat. No. 5,845) as the female parent and an unnamed proprietary seedling identified as UC #3 as the male parent. Hand pollination was performed under greenhouse conditions and seed was germinated under laboratory conditions. Seedlings were individually cultured in a greenhouse before transplanting into a field for evaluation and selection. The varietal denomination of the new variety is `Victoria`. The new variety is a sibling of the variety `De Anza` which is the subject of Plant Patent Application Ser. No. 08/181,860, filed Jan. 13, 1994.
Sustained drought conditions in California have additionally increased the need for improved warm season turf grass varieties. Zoysiagrasses are well adapted for use under such conditions because of their low water use rates and high heat tolerance, however, acceptance has been low primarily due to loss of green coloration during the winter months as such grasses routinely go into dormancy.
`El Toro` Zoysiagrass is a much faster-growing variety of Zoysiagrass than others of the species, and it made Zoysiagrass more attractive to commercial sod producers. However, although `El Toro` exhibits a longer growing season than previously available commercial Zoysiagrass varieties, it does not remain green all year and also tends to turn purple during the onset of winter dormancy. In addition, the leaf texture of `El Toro` is considered to be somewhat coarse. The new variety is an improvement over Zoysiagrass germplasm available for use as turf grass such as `El Toro` Zoysiagrass.
SUMMARY OF THE INVENTIONThe new variety of Zoysiagrass, `Victoria`, exhibits an extended growing season and longer retention of green coloration during the winter season than presently available commercial Zoysiagrass varieties, including `El Toro`. Under moderate climatic conditions such as coastal areas of California, acceptable green color quality can be maintained throughout the year. The new variety is vegetatively propagated from sod, plugs, tillers or stolon and rhizome pieces. Asexual propagation by rhizomes and tillers in Riverside County Calif. has established that the combination of characteristics of the new variety are transmitted through succeeding propagations. Seed stability has not been determined.
Comparison to `El Toro``Victoria` differs from the Zoysiagrass variety `El Toro` in the length of the growing season and in retention of green color during winter. The new variety also has narrower leaves and produces a finer-textured turf than `El Toro` and unlike `El Toro`, it produces substantially no purple pigmentation in leaves, stems or inflorescence.
BRIEF DESCRIPTION OF THE ILLUSTRATIONSThe accompanying illustrations show typical specimens of the vegetative growth of the new variety depicted in color as nearly true as it is reasonably possible to make the same in color illustrations of this character.
FIG. 1 shows the turf appearance of the new variety on Oct. 15, 1993;
FIG. 2 shows the color of the new variety in December 1991;
FIG. 3 shows the color of `El Toro` in December 1991;
FIG. 4 shows the appearance of a single leaf of the new variety as compared to another new variety which is the subject of co-pending U.S. plant patent application Ser. No. 08/181,860, filed Jan. 13, 1994, and `El Toro`;
FIG. 5 depicts the leaf orientation of the same three Zoysiagrass lines identified in FIG. 4; and
FIG. 6 shows stolons of a new variety also compared with the variety of the aforementioned co-pending application and `El Toro` to illustrate the difference in purple pigmentation (anthocyanin) as compared to the variety `El Toro` on Oct. 15, 1993.
DESCRIPTION OF THE NEW VARIETYBecause of its heritage, the new variety `Z88-14` is a hybrid of three Zoysiagrass species, Z. japonica, Z. matrella and Z. tenuifolia. It forms a moderately dense turf that spreads by way of tillers, stolons and rhizomes.
Leaf blades of the new variety are bright green in coloration, near R.H.S. 138A to B, and range from about 1.9 to about 7.0 cm in length, with an average length of about 3.6 cm. The leaf width ranges from about 1.8 to about 3.4 mm with an average leaf width of about 2.7 mm. The shorter leaves are lanceolate in shape while longer leaves become linear. Leaves are rolled in the bud and the ligule is a fringe of hairs. Auricles are absent.
Stolons of `Victoria` are yellowish green to green in color and stolon internode length ranges from about 0.8 to about 3.6 cm with an average internode length of about 2.2 cm. The stem thickness of internodes ranges from about 0.9 to about 1.6 mm with an average thickness of about 1.2 mm.
In its inflorescence, `Victoria` exhibits a spike-like raceme ranging from about 1.6 to about 2.6 cm in length with an average of about 1.9 cm. Spikelets average about 3.0 mm in length with a range of about 2.6 to 3.4 mm. The peduncles are green and the spikelets are yellowish green, becoming straw-colored when mature.
The new variety, like other Zoysiagrasses, is vegetatively propagable from sod, plugs or stolon or rhizome pieces. It performs best in full sun and is best adapted to warmer climates where heat and drought stress occur. Near-freezing night temperatures will reduce color quality, and prolonged exposuure to freezing or near freezing temperatures will induce dormancy. The turf formed by the new cultivar is suitable for situations requiring a durable, high quality lawn that is tolerant of heat and drought stress and is resistant to pests. Such applications would include home lawns, parks and other public planting areas and golf courses.
The accompanying illustrations serve to illustrate the distinctive features of the new variety. As can be seen from FIGS. 1-6, `Victoria`, also referred to as `Z88-14` differs from the other new variety `De Anza` (Z88-8), a sister seedling, compared with `Victoria` and also from `El Toro`. In particular, the absence of purple pigmentation of `Z88-14` is shown in FIG. 5, which is compared with the `El Toro` specimen, which shows extensive purple pigmentation, and the difference in winter color from `El Toro` can be seen in FIGS. 2 and 3.
To further illustrate the difference between the new variety and `El Toro`, tests were conducted at the University of California at Riverside and the University of California South Coast Field Station (SCFS) at Irvine, Calif., to compare the winter color of Zoysiagrasses. The results of these evaluations are shown in Tables I and II. Statistical differences among the cultivars were determined using the Duncan's Multiple Range Test. Values given in Tables I and II that are followed by the same letters are not significantly different from each other at the 5% probability level. For these comparisons, plugs of the grasses were placed on one-foot centers with each replicated three times. Common management practices were used during the 1991-92 period, and after establishment, the grasses were mowed at 3/4-inch height at UC SCPS, and 5/8 inch at UC Riverside. The grasses were irrigated as needed based on evapotranspiration from an on-site CIMIS station at both locations and were fertilized at about 6 pounds nitrogen per 1000 square feet of turf area. Tables I and II present the fall/winter color of the grasses on a regular basis, at both test sites, from November 1992 through March 1993 using a rating system of 1 to 9 with 1 representing dormant turf and 9 representing full green color. Also shown are the soil temperatures in .degree.F. at 4 inches depth for each observation date. Normal soil temperatures at UC SCFS did not go as low as Riverside. Once the temperatures went below 50.degree. F. (at UC Riverside), `El Toro` exhibited considerable color loss while the new variety continued to have acceptable green coloration.
TABLE I
__________________________________________________________________________
OBSERVATION DATE
__________________________________________________________________________
GRASS 10-26-92
11-2-92
11-10-92
11-16-92
11-23-92
11-30-92
12-14-92
12-21-92
12-31-92
__________________________________________________________________________
`El Toro`
7.7 bcde
6.7 cde
7.0 bcd
7.0 abcde
6.3 bcde
6.0 abcde
4.0 defg
4.0 cd
3.3 cd
`Z88-8`
9.0 a
8.0 a
8.3 a
8.3 a
7.7 a
7.7 a
7.3 a
6.7 a
6.3 a
`Z88-14`
8.7 ab
7.7 ab
7.7 abc
7.3 abcd
6.7 abcd
7.7 a
6.3 abc
6.0 ab
5.3 ab
Avg. soil
68 68 63 62 61 58 56 53 54
Temp. (.degree.F.)
__________________________________________________________________________
GRASS 1-4-93
1-20-93
1-25-93
2-1-93
2-8-93
2-16-93
2-22-93
3-1-93
__________________________________________________________________________
`El Toro` 3.0 def
3.3 de
4.0 fg
4.3 de
4.3 e
5.3 efg
6.0 cde
5.7 bcd
`Z88-8` 6.3 a
7.0 ab
7.3 ab
7.7 ab
7.0 ab
8.0 ab
8.0 a
7.7 a
`Z88-14` 5.3 ab
6.0 b
6.0 cd
6.7 bc
5.7 cd
6.3 cde
6.9 abcd
6.5 abc
Avg. soil N/A 56 57 56 57 58 58 55
temp. (.degree.F.)
__________________________________________________________________________
TABLE II
__________________________________________________________________________
OBSERVATION DATE
__________________________________________________________________________
GRASS 11-4-92
11-13-92
11-20-92
11-26-92
12-4-92
12-14-92
12-21-92
12-31-92
__________________________________________________________________________
`El Toro`
9.0 a 8.7 ab
8.0 a 7.3 ab
6.7 bcde
4.7 def
3.0 ef
1.7 ef
`Z88-8`
9.0 a 9.0 a
8.0 a 7.7 ab
8.0 a
6.3 abc
5.7 a
5.7 a
`Z88-14`
8.7 ab
8.7 ab
7.7 ab
7.7 ab
7.7 ab
6.7 ab
5.3 a
5.0 ab
Avg. soil
64 58 58 52 50 52 49 50
temp. (.degree.F.)
__________________________________________________________________________
GRASS 1-8-93
1-13-93
1-20-93
1-25-93
2-5-93
2-12-93
2-17-93
2-24-93
3-3-93
__________________________________________________________________________
`El Toro`
2.0 cd
1.0 d
2.0 ef
2.3 de
2.7 gh
5.0 de
4.7 cd
5.0 ef
6.0 cdef
`Z88-8`
4.3 a
4.3 a
5.7 a
7.0 a
7.0 a
7.0 a
7.0 a
7.3 ab
7.7 a
`Z88-14`
4.3 a
4.3 a
5.3 a
6.7 ab
6.0 bc
6.3 abc
6.7 ab
7.3 ab
7.0 abc
Avg. soil
49 52 53 53 55 56 56 55 55
temp. (.degree.F.)
__________________________________________________________________________
Establishment Rate
Regarding establishment rate, it was found that `El Toro` was the fastest to cover the soil surface with 70% coverage recorded five months following establishment and 100% coverage 10 months following establishment. In comparison, `De Anza`, referred to as Z88-8, provided about 57% coverage after 10 months and `Victoria`, referred to as Z88-14, provided about 34% coverage in the same time period. `De Anza` gave 92% coverage after 10 months and complete coverage 12 months following establishment; `Victoria` gave nearly 60% coverage after 10 months and provided 100% coverage after 13 months. In comparison to other commercially available cultivars, `De Anza` is quicker to establish than `Emerald`, `Belair` and `Meyer`; `Victoria` was equal in establishment rate to `Emerald` but faster than `Belair` and `Meyer`.
Consistently, winter color retention is greatest for `De Anza` and `Victoria` zoysiagrass. As an example, in mid-January during the winter of 1991-1992, on a rating scale of 1-9 with 9 representing deep green color and 1 representing dormant, tan or light-brown color, `De Anza` scored a 5.6, `Victoria` a 6.0 and `El Toro` a 2.2. Of the other commercially available cultivars, `Emerald` scored 2.2 and `Belair` and `Meyer` scored a 1.0.
Other ComparisonsLeaf texture was observed in Summer, 1990 by measuring the leaf blade widths at the collar in mm. `De Anza` and `Victoria` were characterized as having an intermediate blade width (on a scale of coarse, intermediate and fine) with blade widths being about 2.72 and about 2.79 mm respectively. `El Toro` was coarse with an average width of 3.41 mm; `Emerald` was fine textured with a width of 2.25 mm; `Meyer` and `Belair` both coarse textured with widths of 3.29 and 4.23 mm respectively.
The amount of seedhead expression was documented throughout the summer of 1990. `De Anza`, `Victoria` and `El Toro` produced more seedheads than did the commercially available cultivars `Emerald`, `Belair` and `Meyer`. Utilizing a low, medium and high rating system that was based on number of seedheads per square meter, `De Anza`, `Victoria` and `El Toro` were rated high and `Emerald`, `Belair` and `Meyer` were rated low.
All recording of turfgrass quality was also summarized in a low, medium and high category by cultivar. `Victoria` was in the high category as was `Emerald`; `De Anza` and `El Toro` were medium in overall quality ranking; and `Meyer` and `Belair` were in the low rank.
Similarly, `Victoria` had the highest ranking for uniformity of turfgrass sward; `De Anza`, `El Toro` and `Emerald` were of medium ranking; and `Belair` and `Meyer` were rated low.
Leaf blade stiffness was on a 1 to 3 rating system with 1 representing a stiff blade and 3 representing a softer, less stiff blade, `Victoria` was characterized as having a soft leaf blade; El Toro, `Meyer` and `Emerald` were rated intermediate with a score of 2; and `De Anza` and `Belair` were noted to have a stiff blade that characterized the cultivar.
Sod Density and Thatch AccumulationSod density was determined in July, 1994 at the University of California, Riverside Turfgrass Research Facility and at the University of California South Coast Research and Extension Center with density being defined as the number of tillers per unit area and reported on a low to high density scale. `De Anza`, `Victoria`, `El Toro` and `Meyer` were characterized as being of moderate density; `Belair` was low density; and `Emerald` was a high density cultivar. Thatch depth measurements taken at the University of California Turfgrass Research Facility in July, 1994, documented, in mm of thatch depth, that `Emerald` produced the greatest depth of thatch (25.0 mm); `Meyer`, `Belair` and `Victoria` produced an intermediate thatch depth (18.0, 15.0 and 14.0 mm respectively); and `El Toro` and `De Anza` produced the lowest amount of thatch (11.67 and 10.33 mm respectively).
`Victoria` Botanical DescriptionThe grass is a low-growing, perennial, attractive turfgrass that forms a moderately dense sward and spreads by tillers, rhizomes and stolons. The color is identified from The Royal Horticultural Society Colour Chart (R.H.S. Chart) as near 139 C.
Leaf blades are flat and range from about 1.9 to about 7.0 cm in length with an average length of about 3.6 cm. Leaf widths range from about 1.8 to about 3.4 mm with an average width of about 2.7 mm. Shorter leaves are lanceolate in shape, while the longer leaves become linear. Leaves are rolled in the bud. The ligule is ciliate with no hairs near the ligule nor to the sides of the ligule; auricles are absent. Stolon color is near 145 C. Stolon internode length ranges from about 0.8 to about 3.6 cm and averages about 2.2 cm. Stem thickness of the internodes ranges from about 0.9 to about 1.6 mm with an average of about 1.2 mm.
The inflorescence is a spikelike raceme ranging from about 1.6 to about 2.6 cm in length with an average length of about 1.9 cm. Culm height average about 14 cm. Spikelets are one-flowered, appressed against a rachis and on a pedicel that ranges from about 1.3 to about 1.9 mm long with an average length of about 1.7 mm. A spikelet with anthers and stigma exerted have a color of near 150 B; mature spikelets are near 19 B. Spikelets are shiny in both instances. Glumes are sharply pointed by awnless.
It is evident from the foregoing that the new variety retains green coloration for a longer period of time than `El Toro` and is, therefore, more desirable as a turf grass in geographical areas favoring its use.
Claims
1. A new and distinct variety of Zoysiagrass named `Victoria`, substantially as described and illustrated, which has a longer growing season and green color retention in winter than the variety `El Toro`, and substantially no purple pigmentation.
| PP8553 | January 18, 1994 | Staton |
Type: Grant
Filed: Jan 13, 1994
Date of Patent: May 9, 1995
Assignee: The Regents of the University of California (Los Angeles, CA)
Inventors: Victor A. Gibeault (Riverside, CA), Matthew K. Leonard (Romoland, CA), Victor B. Youngner, deceased (late of Riverside, CA)
Primary Examiner: James R. Feyrer
Law Firm: Christie, Parker & Hale
Application Number: 8/180,899
International Classification: A01H 500;
